R. A. Fisher and the Adaptive Landscape:
...My own interpretation is that Fisher was sceptical about the value of the landscape concept as such, because both environmental and genetic conditions were too changeable for the metaphor of a 'landscape' to be useful. For Fisher the question of the 'shape' of the landscape therefore did not arise as a major issue, and he had no need to take a firm view on it. I discuss this interpretation below the fold.
Read the whole thing. Related:R. A. Fisher and Epistasis, Notes on Sewall Wright: Population Size, Notes on Sewall Wright: the Measurement of Kinship, Notes on Sewall Wright: Path Analysis, On Reading Wright and Notes on Sewall Wright: Migration
